Spoiler alert: More research may be needed.
First, a recap. Twenty years ago, Japanese scientist Toru Kishimoto identified polyfunctional thiols as a potential contributor to tropical and other fruity aromas in hops, either on their own or acting synergistically with other hop compounds. He focused on three that are present in passion fruit and sauvignon blanc wine. (For more background, also see “The Complex Case of Thiols,” beerandbrewing.com.)